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  • In genetics complete linkage is defined as the state in which two loci are so close together that alleles of these loci are virtually never separated by crossing over. During sexual reproduction, meiosis occurs during which homologous chromosomes pair, and can exchange sections of DNA. As a result, genes that were originally on the same chromosome can finish up on different chromosomes – genetic recombination. The closer the physical location of two genes on the DNA, the less likely they are to be separated.